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William studied for his BSc at the National University of Ireland, Galway, and graduated in 2003. He followed this with a PhD under the supervision of Prof. William Gallagher at the UCD Conway Institute in Dublin, where his project involved the study of DNA methylation in melanoma cells.
He continued his focus on cancer in his Post Doctoral studies with Prof. Owen Sansom at the CRUK Beatson Institute in Glasgow. During this time he began to work with mouse models of colorectal cancer, primarily focusing on mTOR signaling. In 2016 he became a Junior Group Leader at the NKI.